Berlin-based airline Germania has declared insolvency and ceased operations. 

In a statement Tuesday, the company blamed summer increases in fuel prices, the weakening of the euro versus the dollar, high aircraft maintenance burdens and delays in phasing aircraft into its fleet for the liquidity crunch that has forced the closure. 

Germania operated short- and medium-length commercial and charter flights to destinations in Europe, North Africa and the Middle East. The carrier flew more than 4 million passengers annually, according to its website. 

Germania's closure follows the collapse in late 2017 of what was then Germany's second-largest airline, Air Berlin. U.K.'s Monarch Air also ceased operations in late 2017, and Danish budget carrier Primera Air collapsed last October. 

Germania said that individuals who booked tickets as part of a package deal can contact their tour operator for an alternative flight. 

"For passengers who have booked directly with Germania, there is unfortunately no entitlement to substitute carriage," the company said. 

Germania's sister airlines, Bulgarian Eagle and Switzerland-based Germania Flug, continue to operate.

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